Stuart Hogg will captain the British and Irish Lions for their opening fixture in South Africa, against the Emirates Lions on Saturday.
Hogg was dropped by Exeter for the closing stages of their Premiership title push but the Scotland skipper has been thrust into Warren Gatland’s XV at the first available opportunity.
The 29-year-old full-back only linked up with the squad after Saturday’s Premiership final defeat by Harlequins and has been given an immediate chance to press for Test selection.
Finn Russell starts
He is joined in the backline by Scotland team-mate Finn Russell, who forms an intriguing 10-12 axis with Owen Farrell – one of his rivals for the fly-half duties against the Springboks.
Russell has overcome an ankle injury to make his first appearance of the tour and is picked in a half-back partnership alongside compatriot Ali Price.
A strong Scottish influence runs throughout the team for a fixture that arrives a week after Japan were flattened 28-10 in the pre-tour curtain raiser at Murrayfield.
Hamish Watson, the current Six Nations player of the year, is restored at openside having been forced to withdraw from the back-row to face the Brave Blossoms because of concussion sustained in training.
Chris Harris makes his first appearance of the tour at outside centre, lifting the number of Scotsmen in the starting XV to five. Prop Zander Fagerson is on the bench after recovering from a back spasm.
England dominate the pack where they supply five starters in hooker Jamie George, prop Kyle Sinckler, locks Maro Itoje and Jonny Hill and flanker Courtney Lawes.
It comes after Eddie Jones’ men were overlooked entirely against Japan and this time it is Ireland whose only representation is on the bench.
“It’s good to have arrived in Johannesburg and get the tour under way,” Gatland said. “We had a pleasing win last week up in Edinburgh, but there’s still plenty we need to get right on Saturday.
“I’ve been encouraged by the way the boys have trained this week. I feel it’s gone up a level from where we were in Jersey and we’re starting to see a better understanding of some of our tactics.
“There’s some new player combinations for us to have a look at this Saturday – which is good information for us as the Test series approaches.
“My congratulations to all those playing their first game as a Lion, and also to Hoggy too – who I am sure will lead the side by example.”
British & Irish Lions: 15 Stuart Hogg (c), 14 Louis Rees-Zammit, 13 Chris Harris, 12 Owen Farrell, 11 Josh Adams, 10 Finn Russell, 9 Ali Price, 8 Taulupe Faletau, 7 Hamish Watson, 6 Courtney Lawes, 5 Jonny Hill, 4 Maro Itoje, 3 Kyle Sinckler, 2 Jamie George, 1 Wyn Jones
Replacements: 16 Luke Cowan-Dickie, 17 Mako Vunipola, 18 Zander Fagerson, 19 Iain Henderson, 20 Sam Simmonds, 21 Gareth Davies, 22 Bundee Aki, 23 Elliot Daly
Date: Saturday, July 3
Venue: Emirates Airline Park
Kick-off: 18:00 local (17:00 BST, 16:00 GMT)
Referee: AJ Jacobs
Assistant Referees: Wayne Barnes, Jaco Peyper
TMO: Stuart Berry